Higher Classification: > Kingdom Animalia > Subkingdom Parazoa > Phylum Porifera > Subphylum Cellularia > Class Demospongiae > Subclass Heteroscleromorpha > Order Suberitida > Family Halichondriidae > Genus Ciocalypta

Ciocalypta penicillus Bowerbank, 1862

Rank: Species
Taxon Status: accepted

Relationships towards this taxon

Genus group names

*Ciocalypta*	Bowerbank, 1862	accepted genus name
*Coelocalypta*	Topsent, 1928	synonym
*Leucophloeus*	Carter, 1883	synonym

Species group names

*Spongia cavernosa*	sensu Lamarck, 1814	synonym

Vernaculars (+) (-)

Dutch: penseelspons

English: brush horny sponge; brush sponge sources: National General Status Working Group [Groupe de travail national sur la situation générale]. (2017). Standardized Common Names for Wild Species in Canada [Noms communs normalisés pour les espèces sauvages au Canada].

French: éponge pinceau

German: Pinselschamm

Italian: spugna con granchio facchino

Japanese: アオホテイカイメン

Literature

original description	Bowerbank, J.S. (1862). On the Anatomy and Physiology of the Spongiadae. Part III On the Generic Characters, the Specific Characters, and on the Method of Examination. Philosophical Transactions of the Royal Society. 152(2): 1087-1135, pls LXXII-LXXIV.
basis of record	Ackers, R.G.; Moss, D.; Picton, B.E. (1992). Sponges of the British Isles (‘Sponges V'). A Colour Guide and Working Document. Marine Conservation Society. 1-175.
basis of record	Bowerbank, J.S. (1866). A Monograph of the British Spongiadae. Volume 2. (Ray Society: London): i-xx, 1-388.
additional source	Pouliquen, L. (1972). Les spongiaires des grottes sous-marines de la région de Marseille: Ecologie et systématique. Téthys. 3(4): 717-758.
additional source	Labate, M.; Arena, P. (1964). La fauna dei Poriferi nei laghi di Ganzirri e Faro (Messina). Archivo Zoologico Italiano. 49: 249-280.
additional source	Muller, Y. (2004). Faune et flore du littoral du Nord, du Pas-de-Calais et de la Belgique: inventaire. [Coastal fauna and flora of the Nord, Pas-de-Calais and Belgium: inventory]. Commission Régionale de Biologie Région Nord Pas-de-Calais: France. 307 pp.
additional source	Burton, M. (1956). The sponges of West Africa. Atlantide Report (Scientific Results of the Danish Expedition to the Coasts of Tropical West Africa, 1945-1946, Copenhagen). 4: 111-147.
additional source	Marra, M.V.; Bertolino, M.; Pansini, M.; Giacobbe, S.; Manconi, R.; Pronzato, R. (2016). Long-term turnover of the sponge fauna in Faro Lake (North-East Sicily, Mediterranean Sea). Italian Journal of Zoology. 1-10.
additional source	Santín, A.; Grinyó, J.; Ambroso, S.; Uriz, M.-J.; Gori, A.; Dominguez-Carrió, C.; Gili, J.-M. (2018). Sponge assemblages on the deep Mediterranean continental shelf and slope (Menorca Channel, Western Mediterranean Sea). Deep Sea Research Part I: Oceanographic Research Papers. 131: 75-86.
additional source	Boury-Esnault, N.; Harmelin, J.-G.; Ledoyer, M.; Saldanha, L.; Zibrowius, H. (2001 [2005]). Peuplement benthique des grottes sous-marines de Sagres (Portugal, Atlantique nord-oriental). In: Biscoito, M., A.J. Almeida & P. Ré (Eds). A Tribute to Luiz Saldanha. Boletim do Museu Municipal do Funchal. Sup. 6: 15-38.
additional source	Descatoire, A. (1969). Peuplements sessiles de l'Archipel de Glénan de l'infralittoral rocheux. I. - Inventaire: Spongiaires. Vie et Milieu (B, Océanographie). 20 (1B): 177-209.
additional source	Castric-Fey, A. (1996). Richesse et biodiversité en mer mégatidale: communautés sublittorales rocheuses de la région de Trébeurden-Ploumanac'h (Nord Bretagne France). Cahiers de Biologie marine. 37, 7-31.
additional source	Le Mao, P.; Godet, L.; Fournier, J.; Desroy, D.; Gentil, F.; Thiébaud, E.; Poutinet, L.; Cabioch, L: Retière, C.; Chambers, P. (2020). Atlas de la faune marine invertébrée du golfe Normano-Breton Volume 5 - Autres espèces -. Éditions de la Station biologique de Roscoff, ISBN 82951802971. 308 pp.
additional source	Sarà, M. (1958). Studio sui Poriferi di una grotta di marea del Golfo di Napoli. Archivo Zoologico Italiano. 43: 203-281, pls I-II.
additional source	Burton, M. (1936). The fishery ground near Alexandria. IX. Sponges. Notes and Memoirs of the Fisheries Research Directorate, Cairo. 17: 1-28.
additional source	Burton, M. (1959). Sponges. In: Scientific Reports. John Murray Expedition 1933-34. 10(5). British Museum (Natural History): London. Pp. 151-281.
additional source	Boury-Esnault, N. (1971). Spongiaires de la zone rocheuse de Banyuls-sur-Mer. II. Systématique. Vie et Milieu. 22(2): 287-349.
additional source	Naveiro Millán, A. (2002). Poríferos de la costa da Arrábida (Portugal). Clase Demospongiae. Tesina Unversidad de Santiago de Compostela. 165 pp.
additional source	Borojevic, R.; Cabioch, L.; Lévi, C. (1968). Inventaire de la faune marine de Roscoff. Spongiaires. Éditions de la Station Biologique de Roscoff. pp. 1-44.
additional source	Vacelet, J. (1961). Spongiaires (Démosponges) de la région de Bonifacio (Corse). Recueil des Travaux de la Station marine d'Endoume. 22 (36): 21-45.
additional source	Bowerbank, J.S. (1874). A Monograph of the British Spongiadae. Volume 3. (Ray Society: London): i-xvii, 1-367, pls I-XCII.
additional source	Girard-Descatoire, A.; Castric-Fey, A.; L'Hardy-Halos, M.T. (1995). Inventaire de la Faune et de la Flore sur les fonds rocheux autour de l'île d'Ouessant. Rapport ADMS, Direction Régionale de l'Environnement Bretagne, Conseil Régional de Bretagne, Conseil Général du Finistère, Fonds Européens, Rennes., 148pp. Convention ZNIEFF 94.
additional source	Topsent, E. (1925). Etude des Spongiaires du Golfe de Naples. Archives de Zoologie expérimentale et générale. 63(5): 623-725, pl. VIII.
additional source	Thomas, P.A. (1985). Demospongiae of the Gulf of Mannar and Palk Bay. In: Recent Advances in Marine Biology. New Delhi, James, P.S.B.R. (ed.). Today Tomorrow's Printers and Publishers. 205-365.
additional source	Tanita, S.; Hoshino, T. (1989). The Demospongiae of Sagami Bay. Biological Laboratory, Imperial Household: Japan. i-xiii, 1-197 [in English], pls 1-19; 1-166 [in Japanese], 1 map.
additional source	Topaloğlu, B. (2001). Sponge fauna in the littoral zone of the Marmara Sea. Rapports et Procès Verbaux des Réunions de la Commission Internationale pour l'Exploration de la Mer Méditerranée,. 36, 421.
additional source	Kelly, M.; Edwards, A.R.; Wilkinson, M.R.; Alvarez, B.; Cook, S. de C.; Bergquist, P.R.; Buckeridge, St J.; Campbell, H.J.; Reiswig, H.M.; Valentine, C.; Vacelet, J. (2009). Phylum Porifera: sponges. in: Gordon, D.P. (Ed.) (2009). New Zealand inventory of biodiversity: 1. Kingdom Animalia: Radiata, Lophotrochozoa, Deuterostomia. pp. 23-46.
additional source	Van Soest, R.W.M. (2001). Porifera, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels. 50: 85-103.
additional source	Topsent, E. (1897). Spongiaires de la Baie d'Amboine. (Voyage de MM. M. Bedot et C. Pictet dans l'Archipel Malais). Revue suisse de Zoologie. 4: 421-487, pls 18-21.
additional source	Pulitzer-Finali, G. (1978). Report on a Collection of Sponges from the Bay of Naples. III Hadromerida, Axinellida, Poecilosclerida, Halichondrida, Haplosclerida. Bollettino dei Musei e degli Istituti Biologici dell'Universitá di Genova. 45: 7-89.
additional source	Hiscock, K.; Stone, S.M.K.; George, J.D. (1984). The marine fauna of Lundy. Porifera (sponges): a preliminary study. Report Lundy Field Society. 34: 16-35.
additional source	Hayward, P.J. & J.S. Ryland (Eds.). (1990). The marine fauna of the British Isles and North-West Europe: 1. Introduction and protozoans to arthropods. Clarendon Press: Oxford, UK. 627 pp.
additional source	Van Soest, R.W.M. (1977). Marine and freshwater sponges (Porifera) of the Netherlands. Zoölogische Mededelingen. Rijksmuseum van Natuurlijke Historie te Leiden. 50(16): 261-273.
additional source	Solórzano, M.R. (1990). Poríferos del litoral gallego: estudio faunístico, distribución e inventario. Phd Thesis Unversidad de Santiago de Compostela. 1036 pp.
biology source	Morrow, C.; Picton, B.; Erpenbeck, D.; Boury-Esnault, N.; Maggs, C.; Allcock, A. (2012). Congruence between nuclear and mitochondrial genes in Demospongiae: A new hypothesis for relationships within the G4 clade (Porifera: Demospongiae). Molecular Phylogenetics and Evolution. 62(1): 174-190.

Genbank

External Links

Marine Life Information Network - UK

Citation

de Voogd, N.J.; Alvarez, B.; Boury-Esnault, N.; Cárdenas, P.; Díaz, M.-C.; Dohrmann, M.; Downey, R.; Goodwin, C.; Hajdu, E.; Hooper, J.N.A.; Kelly, M.; Klautau, M.; Lim, S.C.; Manconi, R.; Morrow, C.; Pinheiro, U.; Pisera, A.B.; Ríos, P.; Rützler, K.; Schönberg, C.; Turner, T.; Vacelet, J.; van Soest, R.W.M.; Xavier, J. (2025). World Porifera Database. Ciocalypta penicillus Bowerbank, 1862. Via: Costello, M.J.; Bouchet, P.; Boxshall, G.; Arvantidis, C.; Appeltans, W. (2011) European Register of Marine Species, Accessed through PESI at http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:marinespecies.org:taxname:132575