Higher Classification: > Kingdom Animalia > Subkingdom Eumetazoa > Phylum Chordata > Subphylum Tunicata > Class Ascidiacea > Order Stolidobranchia > Family Styelidae > Genus Styela

Styela clava Herdman, 1881

Rank: Species
Taxon Status: accepted

Relationships towards this taxon

Genus group names

*Styela*	Fleming, 1822	accepted genus name
*Botryorchis*	Huntsman, 1911	synonym

Vernaculars (+) (-)

Dutch: Japanse zakpijp; knotszakpijp sources: Eneman, E. (1995). Knotszakpijp of Japanse zakpijp Styela clava (Herdman, 1882) in de Spuikom van Oostende. [Leathery sea squirt Styela clava (Herdman, 1882) in the Sluice Dock at Ostend]. De Strandvlo. 15(3): 113.;Leewis, R. (2002). Flora en fauna van de zee [Marine flora and fauna]. Veldgids, 16. KNNV Uitgeverij: Utrecht, The Netherlands. ISBN 90-5011-153-X. 320 pp.

English: Asian tunicate; Clubbed tunicate; Eathery sea squirt; folded sea squirt; leathery sea squirt; rough sea squirt sources: Volvenko, I. V.; Orlov, A. M.; Gebruk, A. V.; Katugin, O. N.; Ogorodnikova, A. A.; Vinogradov, G. M.; Maznikova, O. A. (2019). Checklist of the trawl macrofauna from the North Pacific and adjacent seas with information about fishery importance, potential product yield, and price range, supplement to: Volvenko, Igor V; Orlov, Alexei M; Gebruk, Andrey V; Katugin, Oleg N; Ogorodnikova, Alla A; Vinogradov, Georgy M; Maznikova, Olga A (in prep.): Trawl macrofauna of the Far-Eastern Seas and North Pacific: proportion of commercial species, potential product yield, and price rang. Earth System Science Data. PANGAEA - Data Publisher for Earth & Environmental Science.;Leewis, R. (2002). Flora en fauna van de zee [Marine flora and fauna]. Veldgids, 16. KNNV Uitgeverij: Utrecht, The Netherlands. ISBN 90-5011-153-X. 320 pp.;Eneman, E. (1995). Knotszakpijp of Japanse zakpijp Styela clava (Herdman, 1882) in de Spuikom van Oostende. [Leathery sea squirt Styela clava (Herdman, 1882) in the Sluice Dock at Ostend]. De Strandvlo. 15(3): 113.;Hammerson, G.A. (2004).Connecticut Wildlife: Biodiversity, Natural History, and Conservation. ISBN 1584653698

French: ascidie japonaise; ascidie plissée sources: Dewarumez, J.-M.; Gévaert, F.; Massé, C.; Foveau, A.; Desroy, N.; Grulois, D. (2011). Les espèces marines animales et végétales introduites dans le bassin Artois-Picardie . UMR CNRS 8187 LOG/Agence de l'Eau Artois-Picardie: Douai. 138 pp.;Muller, Y. (2004). Faune et flore du littoral du Nord, du Pas-de-Calais et de la Belgique: inventaire. [Coastal fauna and flora of the Nord, Pas-de-Calais and Belgium: inventory]. Commission Régionale de Biologie Région Nord Pas-de-Calais: France. 307 pp.

German: Falten-Ascidie; Ostasiatische Seescheide sources: Leewis, R. (2002). Flora en fauna van de zee [Marine flora and fauna]. Veldgids, 16. KNNV Uitgeverij: Utrecht, The Netherlands. ISBN 90-5011-153-X. 320 pp.;Unterwasser-Welt Ostsee

Japanese: エボヤ sources: Japan Agency for Marine-Earth Science and Technology. (2009 onwards). Biological Information System for Marine Life (BISMaL). Accessed on 2018-11-21.

Russian: Стиела булавовидная

Literature

original description	Herdman, W. A. (1881). Preliminary report on the Tunicata of the Challenger expedition. Cynthiidae. Proceeding of the Royal Society of Edinburgh. 11(3): 52-88.
basis of record	Monniot, C. (2001). Ascidiacea & Sorberacea. In: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels. 50: pp. 352-355.
additional source	Muller, Y. (2004). Faune et flore du littoral du Nord, du Pas-de-Calais et de la Belgique: inventaire. [Coastal fauna and flora of the Nord, Pas-de-Calais and Belgium: inventory]. Commission Régionale de Biologie Région Nord Pas-de-Calais: France. 307 pp.
additional source	Liu, J.Y. [Ruiyu] (ed.). (2008). Checklist of marine biota of China seas. China Science Press. 1267 pp.
additional source	Zenetos, A., S. Gofas, M. Verlaque, M. Cinar, J. Garcia Raso, C. Bianchi, C. Morri, E. Azzurro, M. Bilecenoglu, C. Froglia, I. Siokou, D. Violanti, A. Sfriso, G. San Martin, A. Giangrande, T. Katagan, E. Ballesteros, A. Ramos-Espla, F. Mastrototaro, O. Ocana, A. Zingone, M,. Gambi & N. Streftaris. (2010). Alien species in the Mediterranean Sea by 2010. A contribution to the application of European Union's Marine Strategy Framework Directive (MSFD). Part I. Spatial distribution. Mediterranean Marine Science. 11(2): 381-493.
additional source	Kott, P.; Bradford-Grieve, J.; Esnal, G.; Murdoch, R.C. (2009). Phylum Tunicata: sea squirts, salps, appendicularians, in: Gordon, D.P. (Ed.) (2009). New Zealand inventory of biodiversity: 1. Kingdom Animalia: Radiata, Lophotrochozoa, Deuterostomia. pp. 409-430.
additional source	Izquierdo-Muñoz, A.; Díaz-Valdés, M.; Ramos-Esplá, A.A. (2009). Recent non-indigenous ascidians in the Mediterranean Sea. Aquatic Invasions. 4(1): 59-64.
additional source	Kott, P. (1985). The Australian Ascidiacea part 1, Phlebobranchia and Stolidobranchia. Mem Qd Mus. 23: 1-440.
additional source	Hillock, K. A.; Costello, M. J. (2013). Tolerance of the invasive tunicate Styela clava to air exposure. Biofouling. 29(10): 1181-1187.
additional source	Howson, C.M. & B.E. Picton. (1997). The species directory of the marine fauna and flora of the British Isles and surrounding seas. Ulster Museum Publication, 276. The Ulster Museum: Belfast, UK. ISBN 0-948150-06-8. vi, 508 (+ cd-rom) pp.
additional source	Millar, R. H. (1975). Ascidians from the Indo-West-Pacific region in the zoological museum, Copenhagen (Tunicata, Ascidiacea). Steenstrupia. 3(20): 205-336.
additional source	Zenetos, A., E. Meric, M. Verlaque, P. Galli, C.F. Boudouresque, A. Giangrande, M. Cinar & M. Bilecenoglu. (2008). Additions to the annotated list of marine alien biota in the Mediterranean with special emphasis on Foraminifera and Parasites. Mediterranean Marine Science. 9(1): 119-165.
additional source	van der Land, J. (ed). (2008). UNESCO-IOC Register of Marine Organisms (URMO).
additional source	Streftaris, N., A. Zenetos & E. Papathanassiou. (2005). Globalisation in marine ecosystems: the story of non-indigenous marine species across European seas. Oceanogry and Marine Biology: an Annual Review. 43: 419-453.

Genbank

External Links

Marine Life Information Network - UK

Citation

Shenkar, N.; Gittenberger, A.; Lambert, G.; Rius, M.; Moreira da Rocha, R.; Swalla, B.J.; Turon, X. (2025). Ascidiacea World Database. Styela clava Herdman, 1881. Via: Costello, M.J.; Bouchet, P.; Boxshall, G.; Arvantidis, C.; Appeltans, W. (2011) European Register of Marine Species, Accessed through PESI at http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:marinespecies.org:taxname:103929